Before the ventricular apex is excitated by the electric front, small branches leave the major electrical highways relatively quickly after dividing into a left and a right Tawara branch. These small branches excitate the septum and the papillary muscles. This excitation is pointed towards the heart’s base. The leads pointing in the direction of the electric heart axis—and thus away from this septum and papillary nerve excitation—will initially show a small negative deflection. Here, we refer to the physiological "septal Q wave".
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